Joint Filing Agreement

• Parties: ARES TRADING SA, MERCK SERONO SA COINSINS, SWITZERLAND, AN AFFILIATE OF MERCK KGAA, DARMSTADT, GERMANY, MERCK KGAA DARMSTADT, GERMANY
• Section: Commercial Contracts

Exhibit 99.1

JOINT FILING AGREEMENT

Additional Reporting Person (a): Merck Serono SA

Address: Zone Industrielle

1267 Coinsins, Switzerland

Additional Reporting Person (b): Merck KGaA

Address: Frankfurter Str. 250

64293 Darmstadt, Germany

Designated Filer: Ares Trading SA

Issuer and CUSIP: Intrexon Corporation (46122T102)

Dated: January 7, 2019

ARES TRADING SA ARES TRADING SA

By: /s/ Cédric Hyde By: /s/ Luigia Bocola

Name: Cédric Hyde Name: Luigia Bocola

Title: CFO Title: Finance Manager

MERCK SERONO SA, COINSINS, SWITZERLAND, AN

AFFILIATE OF MERCK KGAA, DARMSTADT, GERMANY

MERCK SERONO SA, COINSINS, SWITZERLAND, AN

AFFILIATE OF MERCK KGAA, DARMSTADT, GERMANY

By: /s/ Cédric Hyde By: /s/ Tearaboth Te

Name: Cédric Hyde Name: Tearaboth Te

Title: CFO Title: Treasury Director

MERCK KGAA, DARMSTADT, GERMANY MERCK KGAA, DARMSTADT, GERMANY

By: /s/ Rando Bruns By: /s/ Tim Nielsen

Name: Rando Bruns Name: Tim Nielsen

Title: Head of Treasury Title: Head of Capital Markets